1) Arrange 2) Catenulate 3) Form into a chain 4) Set up 
to connect as in or by a chain
• (v. t.) To connect, in a series of links or ties; to chain.
<molecular biology> Two or more circular DNA molecules where one or more circles run through the enclosed space of another like links in a chain. ... (16 Mar 1998) ... 
Two or more circular DNA molecules where one or more circles run through the enclosed space of another like links in a chain.
Cat'e·nate transitive verb [ imperfect & past participle Catenated ; present participle & verbal noun Catenating .] [ Latin catenatus , past participle of catenare , from catena chain. See Chain .] To connec...
Type: Term Pronunciation: kat′en-āt Definitions: 1. To connect in a series of links like a chain; for example, two rings of mitochondrial DNA are often catenated.
[v] - arrange in a series of rings or chains, as for spores
catenulate verb arrange in a series of rings or chains, as for spores